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Hino Akane (日野あかね Hino Akane ?) (known as Kelsey in the English dub) A girl from Osaka whose parents run an okonomiyaki shop and is a member of the school's volleyball team.

Her catchphrase is "Ultra Happy!" She transforms into Cure Happy( Glitter Lucky in the dub) and uses the power of light. Like many of the lead Cures before her, Miyuki is cheery and optimistic, but also something of a klutz.

Hoshizora Miyuki (星空みゆき Hoshizora Miyuki ?) (known as Emily in the English dub) A fourteen-year-old girl with an interest in picture books and fairy tales, who transferred to Nanairogaoka Middle School.

It was 3 years scene the ending of Smile Pretty Cure!,the cures meets 2 new girls who are going to attend there old school,however the Bad End Kingdom came back for its revenge,the cures plus the new girls got the power to become pretty cure again and will stop them